| ORRIS | Violet-scented rootstock of the Iris florentina and related species; or, a form of gold or silver lace (5) |
| SATYRID | Butterfly of a group that includes the browns, heaths, ringlets, and related species (7) |
| EBONY | Hard dark wood of the Indian tree Diospyros ebenum and related species (5) |
| AURIC | Relating to a form of gold |
| ORMOLU | A form of gold leaf prepared for gilding bronze (6) |
| TUBER | In botany, the genus of truffles; the thickened subterranean stem of the Jerusalem artichoke, potato or yam; or, the rootstock of the cyclamen or the dahlia (5) |
| TWITCH | A subduing twist of a horse's lip with a noose of the same name; a supposed nervous jerk, jolt, sudden start or proverbial feather-ruffling flutter of a birder when spotting a rare species; or, a pang |
| RING | A band of gold or silver, sometimes set with gems; an "elf circle" of champignon mushrooms on a woodland floor; one of the concentric markings in the trunk of a tree; or, a disc of matter orbiting Sat |
| BEAM | Word for a tree originally, hence its survival in names of arbors including the "horn" and "white" species; or, a transverse timber of a ship, hence the greatest width of said vessel (4) |
| PARAGON | Word for a touchstone for testing the purity of gold or silver originally, later a perfect example or gold standard; or, a flawless diamond (7) |
| WALK | Group of snails or wagtails; a tree-lined avenue; or, a form of perambulation taken as a leisurely ramble or competitive foot-race (4) |
| PURL | One of the basic knitting stitches; or, a form of coiled silver or gilt thread used in goldwork (4) |
| HEAT | Spicy quality of a chilli; preliminary eliminating round in a race; or, a form of thermal or kinetic energy (4) |
| GROOVE | A fissure, flute or other furrow; good feeling or fine fettle; a fixed routine; or, a form of funk music (6) |
| TISSUE | Word for a rich cloth of gold or silver threads first, later cellular matter of an animal or a vegetable; or, a hankie (6) |
| RALLY | Sequence of point-deciding to-and-fro shots in tennis, squash or badminton; mass gathering for a cause; or a form of motorsport (5) |
| MORSE | Colin Dexter's inspector character; or, a form of code based on a series of dots and dashes (5) |
| SPINNING | Process of converting wool, silk, cotton or flax into yarn; or, a form of cardio on a stationary bike (8) |
| ESCAPE | Act of breaking free; avoidance of danger or harm; or, a form of distraction from reality/routine (6) |
| GUILLEMOT | Auk forming a "bazaar" with its own species or a "seabird city" with a myriad of others including puffins and kittiwakes (9) |
