| ARVICOLA | The water-rat genus of voles (8) |
| RESOLVE | Determination about distribution of voles (7) |
| COYPU | McCoy put out by the water rat (5) |
| VOLE | Not in favour of Leo tormenting the water rat |
| OTTER | The Water Rat's friend |
| HOLLOWAY | Colin Friels in Water Rats, Frank ... (8) |
| CHRISTEY | Steve Bisley in Water Rats, Jack ... (8) |
| TRUFFAUT | Francois, director of the films Fahrenheit 451 and Baisers voles (8) |
| RODENTIA | Order of mammals including beavers and water voles (8) |
| DEESMART | Played Alex St Clare in Water Rats (3,5) |
| WETLANDS | Bogs, fens, marshes, reed beds and other freshwater habitats whose species include dippers, herons, kingfishers, otters and water voles (8) |
| DISSOLVE | Woman's voles unexpectedly vanish (8) |
| OVERSOLD | Muddled red voles promoted too strongly (8) |
| WATERRAT | Mistaken name of a riverbank or "riparian" vole, which is an expert diver and swimmer, widely associated with The Wind in the Willows by extension, a boatman, pirate or sailor; or, a person who is fon |
| MUSQUASH | Large aquatic American animal akin to the vole (8) |
| LOVESICK | Pining vole turning poorly (8) |
| OVERLOOK | Fail to notice rook and vole looking distressed (8) |
| DOVEDALE | Derbyshire river valley (*dead vole) (8) |
| ALOEVERA | Vole originally entering wild area for juicy plant |
| LOOMOVER | Vaguely threaten with a moor vole gone mad (4,4) |
