| GOWER | The -; Welsh peninsula designated Britain's first AONB (5) |
| LLEYN | Welsh peninsula - in valley (Nene) (5) |
| STOUR | River flowing through the Dedham Vale AONB and depicted in John Constable's The Hay Wain (5) |
| CHASE | Cranborne -; AONB overlapping the boundaries of Dorset, Hampshire Somerset and Wiltshire (5) |
| ESSEX | A home county, site of Audley End House, parts of the Dedham Vale AONB and Sir Alfred Munnings' former home Castle House (5) |
| LARNE | Learn about ferry port in Antrim Coast and Glens AONB (5) |
| BARRA | Hebridean island linked to Vatersay - in AONB, Arran (5) |
| BORTH | Resort on Ceredigion Coast path... in AONB or thereabouts (5) |
| BACON | The medieval scholar Roger ____ is often described as Britain's first scientist |
| MINOR | Which Morris model became Britain's first million-selling car? (5) |
| CAROL | Journalist whose mother was Britain's first female Prime Minister, _ Thatcher (5) |
| PERCY | Britain's first celebrity TV gardener, ___ Thrower (5) |
| LUCAS | Caroline ___, Britain's first Green MP (5) |
| ASTOR | Nancy, Britain's first lady elected as MP |
| RIPON | Cathedral city in North Yorkshire whose racecourse hosted Britain's first horse race for female jock |
| GLOWER | Lost at first in Welsh peninsula, scowl angrily |
| COOLINGTOWER | Power station feature, stylish in Welsh peninsula, limiting temperature (7,5) |
| GOWEST | Saint on Welsh peninsula denied right to die |
| WYE | Valley and AONB on the English/ Welsh borders, site of Tintern Abbey and a river of the same name (3) |
| PENNINES | Forming part of the second-largest AONB after the Cotswolds, a range of hills nicknamed the "backbone of England" (8) |
