| SPIRE | Steeple; blade of grass; apex of a shell such as that of the periwinkle; or, a towering stem of a flower such as a delphinium or foxglove (5) |
| CREEPER | A plant such as periwinkle, or a tree-hugging bird (7) |
| NUT | Botanical seed within a shell such as an acorn, filbert or conker; or, a culinary word for a fruit or kernel such as a pistachio, almond, cashew or macadamia (3) |
| MOLLUSC | An animal with a fold of skin secreting a shell such as an oyster (7) |
| VINCA | Any plant of the periwinkle genus (5) |
| WIN | Allure of the periwinkle (3) |
| RACHIS | Whether referring to a backbone, a blade of grass, a feather shaft, a floral stem or the axis of a compound leaf, Greek "spine" is the unifying term (6) |
| SPEAR | A leister, pike, trident or other such pointed javelin; a blade of grass; a stem of asparagus; or, a reed (5) |
| COMPOSITE | Botanical description of a flower of a plant in the aster family such as that of the daisy, dandelion or chamomile (9) |
| PIE | Filled pastry shell such as that in the stargazey style or one baked with a ceramic "bird" in its top crust (3) |
| RIDDLE | A dark or puzzling utterance or question, such as that of the Sphinx; an enigma; or, a coarse-meshed sieve (6) |
| CRUSTA | Word for a hard coating, layer or shell, such as the crystallised lip of a sugar-dipped cocktail glass (6) |
| MIGRATION | Seasonal journey of the population of a species from one habitat to another, such as that of the humpback whale, Atlantic salmon, woodcock, swift or painted lady butterfly (9) |
| BRACT | From the Latin meaning "thin plate of metal, gold-leaf", a modified often-colourful leaf such as that of the red or white poinsettia or "Christmas rose" (5) |
| MUSSEL | "Clappy-doo" with a mother-of-pearl-lined shell, often found anchored to a rock pool with other sea creatures such as an anemone, limpet, periwinkle or starfish (6) |
| SOLIDANGLE | Intersection of three or more planes, resembling a corner or the apex of a cone (5,5) |
| SPIRES | Old English word for tall slender stems or blades of grass that came to mean long tapering objects, such as the tops of church steeples (6) |
| DRAM | A word for 1/8 of a fl oz, hence a small drink, nip or tot of whisky (4) |
| CHOREOLOGY | The study and notation of ballet and dance movement, such as that of the Benesh discipline for example (10) |
| CRUSTACEAN | Arthropod with a hard shell, such as a lobster, crab, shrimp or woodlouse (10) |
