| DUNLIN | Small wading bird with a slightly down-curved bill that breeds in the uplands of Scotland, Wales and England |
| MOUNTAINWAGTAIL | Slender, insect-eating songbird Motacilla clara, found in the uplands of sub-Saharan Africa (8,7) |
| SABINES | ___ gull is a small seabird that breeds in the Arctic (7) |
| BRITISHLIONS | Touring rugby-union team of Scotland, Wales and England (7,5) |
| ROXBURGHSHIRE | Former county in Southern Uplands of Scotland (13) |
| HILLY | Was Henry out of sorts, by the end in the uplands? (5) |
| BIRD | The great skua, or bonxie, is the only ____ that breeds both in the Arctic and in the Antarctic (4) |
| SANDPIPER | Small wading bird with a slender bill (9) |
| PTARMIGAN | Bird of the grouse family which breeds in the mountains of the Highlands (9) |
| RUGBY | Contested at the Six Nations Championship between England, Ireland, Scotland, Wales, Italy and France, one of the sports or games that originated in the UK (5) |
| SANDMARTIN | Bird of the swallow family that breeds in riverbanks and quarries (4,6) |
| RAIL | Small wading bird with short wings, long legs and dark plumage (4) |
| FIRSTMINISTER | Title of heads of government in Scotland, Wales and Northern Ireland (5,8) |
| DORSET | Prehistoric peoples were active on the chalk uplands of this area. There are abundant monuments from the Neolithic Period, Bronze Age, and Iron Age. One monument, Maiden Castle, was the main town in R |
| DOTTEREL | A small wading bird of the plover family (8) |
| STINT | Word for an allowance; an allotted period of work; or, a little wading bird, sometimes called a peep (5) |
| BARNACLE | ____ goose, medium-sized one that breeds in Greenland and the Russian Arctic (8) |
| UNITEDKINGDOM | A monarchy made up of England, Scotland, Wales and Northern Ireland (6,7) |
| KNOT | Small wading bird of the sandpiper family (4) |
| WATERPIPIT | Short, stocky bird, Anthus spinoletta, which breeds in the mountains of southern Europe (5,5) |
