| CALAMUS | Quill of a feather |
| BARREL | Item made by a cooper; belly of a horse; or, the quill of a feather (6) |
| SPINE | Heartwood of a tree; a quill of a hedgehog; a bound edge of a book; a series of vertebrae; or, courage (5) |
| TWO | Number of days before the quills of a newborn porcupine hardens |
| SPINES | The quills of a hedgehog (6) |
| TRUNDLE | A small wheel or castor; the act of rolling slowly; a wheeled bed; a quill of gold thread in heraldry; or, a hoop (7) |
| HUMAN | Like Peter Quill of the Guardians of the Galaxy, in part |
| FLOATS | Buoyant things, such as the balls of ballcocks, corks/quills of fishing-lines, dolphins for mooring, lifebelts, rafts or scoops of ice cream in cola (6) |
| SHAFT | An arrow or spear; a sunbeam, lightning bolt or other column of light; the rachis or rib of a feather; one of a pair of thills of a carriage or cart; or, a well-like passage into a mine (5) |
| RAMUS | Latin word for "branch", thus for a branching of a nerve; a barb of a feather; a process/arm of a bone such as the mandible or pubis; or, a twig (5) |
| BARB | A jag of an arrow or a fish-hook; a beard-like filament of a cyprinid; a harl or ramus of a feather's vane; or, a pointed or wounding remark (4) |
| STICKS | Quills of cinnamon bark; bows for fiddles; supports for candles; or, hurdles in steeplechasing (6) |
| VANE | Flat part on either side of a feather; a sail of a windmill; or, a short word for a weathercock for indicating the wind's direction (4) |
| WHITE | Colour of a feather used as a mark of perceived cowardice or of a flag symbolising truce or surrender (5) |
| VEXILLUM | Vane of a feather; or, a type of ancient Roman military standard (8) |
| GRAN | Maurice, English TV scriptwriter and one half of a duo that created sitcom Birds of a Feather (4) |
| HERL | A barb or filament of a feather used to dress a fishing fly |
| BIRDS | Word that when followed by Of A Feather gives the title of a classic sitcom (5) |
| LINDA | Star of Birds of a Feather, ----- Robson (5) |
| PAULINE | _ Quirke, star of Birds Of A Feather (7) |