| GALTON | Polymath cousin of Charles Darwin who coined the alliterative phrase "nature versus nurture" and was the first to show how fingerprints could be used to identify individuals (6) |
| TENNYSON | Famous Victorian poet who gave us the phrase: 'Nature, red in tooth and claw' (8) |
| FRANCISGALTON | Relative of Charles Darwin who devised the first weather map |
| NATURE | ___ versus nurture |
| MARINA | Second cousin of Charles III, _ Ogilvy (6) |
| BLOODBATH | Slaughter of the alliterative? (5,4) |
| ROEDEAN | Brighton's iconic cliff-top "Eton for girls", founded by three sisters, Dorothy, Millicent and Penelope, to nurture and cultivate young ladies as if precious pearls (7) |
| REAR | Nurture and bring up to maturity / back or hind part |
| REARGUARD | Nurture and protect tail enders |
| SHREWSBURY | With over 600 listed buildings, the county town of Shropshire, birthplace of Charles Darwin (10) |
| EVOLUTION | From the Latin meaning "unrolling", a process described by Charles Darwin in his theory of natural selection; or, one of a series of military manoeuvres (9) |
| ORIGIN | "The ___ of Species" (monumental work of naturalist Charles Darwin (1809-82)) |
| ELMOSFIRE | Electrical weather phenomenon St ..., noted by Charles Darwin as pointing the masts of the HMS Beagl |
| GALAPAGOS | During his voyage on the Beagle, Charles Darwin studied the vast number of species on this archipela |
| BEAGLE | Name the ship in which Charles Darwin was the official scientist (6) |
| TURNIPS | What vegetable was the subject of notes made by Charles Darwin on his wedding night? (7) |
| INSTINCT | "The very essence of _ is that it's followed independently of reason": Charles Darwin (8) |
| MEADOWS | Hayfields such as those surveyed by Charles Darwin at his former home Down House, where he developed the theories in On the Origin of Species (7) |
| ERASMUS | Grandfather of English naturalist Charles Darwin, noted as a physician and a poet (1731-1802) (7) |
| HMSBEAGLE | On which surveying ship was Charles Darwin the official scientist? (3,6) |
