| RODENTIA | Order of mammals including beavers and water voles (8) |
| PRIMATES | An order of mammals including man and monkeys (8) |
| MURID | Any rodent of the largest family of rodents (and of mammals), including the rats, mice, voles etc. (5) |
| WETLANDS | Bogs, fens, marshes, reed beds and other freshwater habitats whose species include dippers, herons, kingfishers, otters and water voles (8) |
| CETACEAN | Relating to an order of aquatic mammals, including sperm whales, dolphins, etc (8) |
| EDENTATE | Order of mammals which includes the anteaters, sloths and armadillos (8) |
| EDENTATA | Order of mammals which includes the anteaters and sloths (8) |
| MONOGAMY | What is practised by gibbons, swans, Eurasian beavers and bald eagles? (8) |
| TRUFFAUT | Francois, director of the films Fahrenheit 451 and Baisers voles (8) |
| ARVICOLA | The water-rat genus of voles (8) |
| RIVERS | Watercourses whose riparian wildlife includes kingfishers, herons, otters and water voles (6) |
| SCUTTLES | Word for the swift scurries of mice, moles, rats or voles; hatchways; trapdoors; or, from "dishes, platters, trays", fireside buckets for coal (8) |
| DISSOLVE | Woman's voles unexpectedly vanish (8) |
| OVERSOLD | Muddled red voles promoted too strongly (8) |
| TWELVE | A dozen water voles soar away (6) |
| SIRENIA | An order of aquatic mammals, including the dugong and the manatee (7) |
| RATTY | Water vole's name in The Wind in the Willows (5) |
| PEEVE | Bug water vole's eviscerated |
| CETACEA | Order of aquatic mammals, including dolphins (7) |
| INSECTIVORA | Order of mammals that includes hedgehogs, moles and shrews (11) |