| ORIGIN | On the ___ of Species, work on evolution by Charles Darwin published in 1859 (6) |
| ERNST | Leading twentieth-century biologist, ___ Mayr, who furthered Darwin's work on evolution and genetics |
| THESELFISHGENE | Book on evolution by Richard Dawkins (3,7,4) |
| ALFRED | _ Bussel Wallace formulated his theory of evolution by natural selection independently of, and before publication by, Darwin (6) |
| DARWIN | Naturalist whose findings collected on his five-year Beagle voyage formed the basis of his theory of evolution by natural selection (6) |
| IGUANA | Large Galapagos lizard - they were described by Charles Darwin as ugly animals with a singularly stu |
| MEADOWS | Hayfields such as those surveyed by Charles Darwin at his former home Down House, where he developed the theories in On the Origin of Species (7) |
| EVOLUTION | From the Latin meaning "unrolling", a process described by Charles Darwin in his theory of natural selection; or, one of a series of military manoeuvres (9) |
| SPECIES | 1859 book by Charles Darwin, On The Origin of ... |
| ATALEOFTWOCITIES | Novel by Charles Dickens published in 1859 (1,4,2,3,6) |
| ADAM | - - - Bede, George Eliot's first novel, published in 1859, set in the community of Hayslope (4) |
| BEDE | Adam ___, George Eliot's first novel, published in 1859; set in the community of Hayslope (4) |
| ELMOSFIRE | Electrical weather phenomenon St ..., noted by Charles Darwin as pointing the masts of the HMS Beagl |
| TURNIPS | What vegetable was the subject of notes made by Charles Darwin on his wedding night? (7) |
| GALAPAGOS | - Islands, in the Eastern Pacific known for their endemic species studied by Charles Darwin (9) |
| CHARLESDARWIN | English naturalist who formulated the theory of evolution by natural selection (7,6) |
| OBLOMOV | The best known novel by Russian writer Ivan Goncharov, first published in 1859 |
| RICHARD | First great novel of George Meredith, published in 1859 (3,6,2,7,7) |
| THE | First great novel of George Meredith, published in 1859 (3,6,2,7,7) |
| FEVEREL | First great novel of George Meredith, published in 1859 (3,6,2,7,7) |
