| NATUREBOOK | Make up title on the origin of species (6,4) |
| DARWIN | Josiah Wedgwood's grandson who was the naturalist on the five-year Beagle voyage and later published his theory of evolution in On the Origin of Species (6) |
| SMATTERING | Superficial knowledge of The Origin of Species making a difference? (10) |
| ERASMUSDARWIN | What name was shared by the grandfather and brother, both noted scientists, of the naturalist who published On the Origin of Species (1859) |
| CHARLES | The Prince of Wales or On The Origin Of Species' Mr Darwin |
| PSEUDONYMS | My opus ends unexpectedly with made-up titles (10) |
| NSELECTION | "The Origin of Species" concept |
| THEORY | Darwin's On The Origin Of Species put forward natural selection as the mechanism in his ... of evolu |
| DARWINISM | Theory of biological evolution first formulated in the 1859 work On the Origin of Species (9) |
| CHARLESDARWIN | English naturalist, author of On The Origin Of Species (7,6) |
| BEAGLE | Ship on which Darwin collected material for "On the Origin of Species" |
| NATURAL | - Selection; Charles Darwin's theory of evolution in On the Origin of Species (7) |
| MEADOWS | Hayfields such as those surveyed by Charles Darwin at his former home Down House, where he developed the theories in On the Origin of Species (7) |
| FACTIOUSBOOK | "On the Origin of Species," for one? |
| SNOOPY | Beagle: Journey's End following On the Origin of Species, revolutionary work (6) |
| FINCH | Bird studied in Darwin's "On the Origin of Species" |
| EVOLUTION | Development and change studied in phylogeny and discussed in Charles Darwin's On the Origin of Species or Yuval Noah Harari's Sapiens (9) |
| SCREED | Long piece of writing on the origin of some belief system (6) |
| AIR | 2023 film based on the origin of a Nike shoe brand in the 1980s |
| SPECIES | Book by Charles Darwin, On The Origin of ... |
