| PAVLOV | Ivan ___, Russian physiologist noted for his study of conditioned reflexes in dogs; Nobel Prize for Physiology or Medicine (1904) |
| IVAN | _ _ _ _ Pavlov, Russian physiologist noted for his work on conditioned reflexes (4) |
| HAVELOCK | ____ Ellis, physician and social reformer born in 1859 noted for his study of human sexuality (8) |
| SWAMMERDAM | Dutch naturalist, biologist and microscopist noted for his study on bees and for being the first per |
| ETTY | British painter, best known for his study of the nude (4) |
| IVANPAVLOV | Which Russian physiologist observed conditioned salivary responses in dogs? (4,6) |
| TURGENEV | Ivan ___, Russian author of the novels Rudin and Fathers and Sons (8) |
| LOTTO | Venetian painter whose Portrait of a Young Man in his Study includes a detail of scattered rose petals thought to allude to the transience of love; or, bingo, a draw or housey-housey (5) |
| STTUDY | This leading thespian can always be seen sitting in his study, admiring the view of his Cornish village (2,4) |
| RUSKIN | Victorian art critic whose teaching collection (including his study of a kingfisher) can be viewed at the Oxford University Ashmolean Museum (6) |
| GONCHAROV | Ivan ___, Russian novelist best known as the author of Oblomov |
| INTRAYS | At the back of his study, Matt from St Martin has an arrangement of containers full of paperwork that needs attending to (2-5) |
| UFOLOGIST | He'll want to get shot of visitors from afar in his study |
| EXERTONESELF | Make physical effort to note reflexes in a poor state (5,7) |
| LYTTA | Fibrous, muscular band of cartilage located on the underside of the tongue in dogs (5) |
| ORNITHOLOGIST | Scientific type with a lot of fliers in his study (13) |
| DEMON | He has a fiendish way of working in his study (5) |
| DURER | Albrecht ___ (1471-1528), foremost artist of the German Renaissance; Saint Jerome in his Study (engraving, 1514), The Four Apostles (painting, 1526) etc. (5) |
| MICHAELFARADAY | Einstein kept a picture of him in his study |
| ITHCHYOLOGIST | With histology itch, is he angling to become specialised in his study (13) |