| SPINE | Heartwood of a tree; a quill of a hedgehog; a bound edge of a book; a series of vertebrae; or, courage (5) |
| QUILL | Spine or prickle of a hedgehog; a roll of cinnamon bark; or, a feather as a pen (5) |
| LEMONYSNICKET | Pen name of US children's writer Daniel Handler, author of the 13-book A Series of Unfortunate Events (1999-2006) (6,7) |
| TRUNDLE | A small wheel or castor; the act of rolling slowly; a wheeled bed; a quill of gold thread in heraldry; or, a hoop (7) |
| BACK | One's dorsum containing a series of vertebrae synonymous with courage or strength of character (4) |
| SPINES | The quills of a hedgehog (6) |
| ACORN | A fruit of a tree, a thing used to make syrup (5) |
| FILLET | From "thread", a word for a narrow ribbon tied round the head originally, later a boned side of fish or slice of meat; a decorative line impressed on the cover of a book; a narrow piece of wood; or, a |
| LEAF | A folio of a book; a botanical structure known in its embryonic form as a cotyledon; or, gold in the form of foil, used in gilding (4) |
| JACKET | Skin of a potato; dust cover of a book; a record sleeve; or, an outer garment (6) |
| ROCKET | From eruca "caterpillar, hairy plant", thus related to "hedgehog", a garden salad plant with a fiery-sounding name and peppery or spicy-tasting leaves that are somewhat jagged, like a cog (6) |
| ECHINO | From the Greek for "sea urchin, hedgehog", a word used in combining form to indicate prickly or spiny (6) |
| SWATCH | Small sample of carpet, cloth or wallpaper; a number of said testers bound as a book; a computer's palette of active colours; or, a spot of ink for matching during printing (6) |
| PLUCK | A short sharp pull of an eyebrow, fowl's feather or fruit from a tree; a strum or twang of a musical string; or, the heart, liver and lungs of an animal, hence courage, guts, heart or spirit (5) |
| CROWN | Another name for a diadem; the upper part of a hat, cut gem or a tree; a clasping metal cap for a bottle; or, a stag's surroyals collectively (5) |
| PASSAGE | An alley; an extract from a book; a journey by air/sea; a section of a piece of music; or, the advance of time (7) |
| BARREL | Item made by a cooper; belly of a horse; or, the quill of a feather (6) |
| ARRAY | Impressive display or collection; rich clothing; a group of hedgehogs; a panel of jurors; or, a matrix (5) |
| THUMB | Series of notches on the outer edge of a book's pages marking the various sections; ... index |
| SHEAF | Archer's set of typically 24 arrows carried in a quiver; or, a bound bundle of harvested wheat, rye or corn stalks (5) |
