| ALLOA | Former burgh of barony in the Central Lowlands (pop about 21,000), once a major port (5) |
| ROSEHEARTY | Old fishing village in the north-east, created a burgh of barony in the 1680s (10) |
| STONEHAVEN | Town in the north-east (pop about 12,000), a former burgh of barony and former county town (10) |
| LEEK | English market town in the West Midlands(pop about 21,000) (4) |
| SALCOMBE | Resort in south Devon that was once a major port for the fruit trade (8) |
| IWOJIMA | In 1945, about 21,000 Japanese and 6800 Americans were killed in the capture of which tiny island? (3,4) |
| DUNS | Former county town in the Borders (pop about 2,500), created a burgh of barony in 1490 (4) |
| NAIRN | Seaside resort in the Highlands (pop about 10,000), once a major fishing port (5) |
| STRATHAVEN | Town in south Lanarkshire (pop about 7,500), created a burgh of barony in 1541 (10) |
| AIRDRIE | Large town near Glasgow (pop about 37,000), created a burgh of barony In 1821 (7) |
| ADANI | Indian conglomerate that signed a deal for a major port development in Sri Lanka. |
| LINEN | Fabric, once a major Scottish export (5) |
| THURSO | Town in Highland Region (pop about 8,000), a burgh of barony from 1633 (6) |
| ALLOW | Permit to enter Central Lowlands (5) |
| ELGIN | Royal burgh of Scotland |
| MOTHERWELL | Former burgh in Lanarkshire (pop 32,000), once a centre of steel production (10) |
| ABERDARE | Town in south Wales (pop about 30,000), once a major coal-mining centre (8) |
| SPARTA | Small town in southern Greece (pop about 17,000), once a major military power (6) |
| BREMEN | Commercial and industrial city in NW Germany with a major port on the River Weser (6) |
| ULTRARUNNING | Sport in which a road race from London to Brighton was once a major event |
