| STROKES | Flourishes or marks made by paintbrushes, pencils or pens; or, the rhythm-setting oars/rowers seated nearest to the sterns of shells (7) |
| ROLLS | Buns; furled cylinders of cake, carpet, cloth, newspaper, parchment etc; strips of material with pockets for paintbrushes, pencils or tools; lists of names; trills; or, throws of dice (5) |
| FOLD | Herd of Highland cattle; a sheep pen; or, the division between the top and bottom halves of a broadsheet (4) |
| ASHORE | He's setting oar on dry land (6) |
| ROLL | A cylinder of gift-wrap or other paper; a strip of cloth or leather with pockets for paintbrushes, pencils, sewing kits or tools; or, a type of brass gilding wheel used in bookbinding (4) |
| DRAWINGS | Pictures made with pencils or pens (8) |
| CURLICUES | Intricate ornamental flourishes or twists, carved or made with a pen (9) |
| GROOVE | A track cut into a record for a stylus; or, the rhythm in jazz, funk or soul (6) |
| METRE | Length unit or the rhythm of a poem |
| LINERS | Fine paintbrushes or kohl pencils for contouring or defining; planes or ships that are part of commercial fleets regularly flying/plying between places; or, the facings of garments (6) |
| CRAYONISTS | Term for artists or scratchboarders who colour in, draw or illustrate with contes, pastels or other pencils or sticks of chalk, pipeclay or wax (10) |
| MEERKATS | Tee marks made by the mongooses (8) |
| FEATHERS | Plumes or quills traditionally fitted to arrows by fletchers or used as pens; or, long tufts of hair on the fetlocks of heavy horses (8) |
| SCRIBES | Ancient writers or copyists whose quills or "pens" recorded history long before the printing press (7) |
| PLIMSOLLMARKS | What are the lines or marks on the hull of British merchant ships that indicate the depth they may be submerged during loading? (8,5) |
| TICKS | Marks made by arachnids (5) |
| SCORER | He counts marks made by kitchen utensil on prime stead (6) |
| BLOTS | Marks made by bolts (5) |
| IMPRINTS | Marks made by pressure |
| WEALS | Marks made by whipping (5) |
