| ARIL | Botanical structure such as the mace surrounding nutmeg, ruby "jewel" of a pomegranate pip or the scarlet "berry" cup of a yew seed (4) |
| TREY | Domino or playing card with three pips; or, the third tine of a deer's antler (4) |
| DONTMAKEWAXES | Bad advice for the SC Johnson company? |
| BITTER | German marque behind cars like the SC and CD (6) |
| PEARL | Ring seating tip of ruby jewel (5) |
| FRUIT | Nutrient-rich botanical structure such as a berry or a melon (5) |
| SEED | Botanical structure such as a peanut or almond |
| LEAF | A folio of a book; a botanical structure known in its embryonic form as a cotyledon; or, gold in the form of foil, used in gilding (4) |
| MONUMENT | From the Latin meaning "remind", a commemorative structure such as the Cenotaph in Whitehall; or, a building of historical importance (8) |
| DOMINO | One of a set of 28 rectangular gaming tiles bearing one to six pips; or, a hooded cloak worn for a masquerade ball with a half-mask of the same name (6) |
| SEEDPEARLS | A mollusc's nacre gifts, no bigger than pips or grains of sand, traditionally adorning a Victorian jewel, such as a brooch or an eternity band (4,6) |
| CHIASMA | Crossing over of two parts or structures such as the fibres of the optic nerves in the brain |
| CHATON | French word for a catkin, a kitten or for the bezel, gemstone or foil-backed jewel of a ring |
| STEMS | Botanical structures supporting flowers, fruits, leaves or the caps of mushrooms; vertical strokes in letters or musical notes; or, the main lines of descent in families (5) |
| STROBILUS | Technical term for the cone of a coniferous plant, extended to cone-shaped structures such as the fruit of the hop (9) |
| CYMBIFORM | Greek-derived word for something shaped like a "boat, keel, ship", from a bone, gravy jug and incense navicula to a botanical structure that is hollowed or curved like a little skiff (9) |
| DOPE | From "sauce, dip", a narcotic drug, originally from the opium poppy seed or pip; or, a word for "cool, great" (4) |
| DAM | Structure such as the Aswan in Egypt and the Three Gorges in China |
| QUATRE | French for a playing card, domino or side of a die with four pips; or, a ballet sequence for four dancers (6) |
| TOWER | Structure such as the Shard, Gherkin, Space Needle or Long Ta (5) |
