| SHELLER | Beachcomber or conchologist who collects the exoskeletons of cockles, cowries, periwinkles, scallops, wentletraps and other molluscs (7) |
| SEASHELL | Sometimes emulated as a chocolate or praline, an object such as a cockle or scallop, collected by a beachcomber or conchologist (8) |
| SCLERITE | Hardened body part, especially one of the plates making up the exoskeleton of an arthropod (8) |
| ECDYSIS | Shedding, as of old skin in reptiles or the exoskeleton in insects and crustaceans (7) |
| EPICUTICLE | Waxy outermost layer of the exoskeleton (integument) of insects, typically impermeable (10) |
| TANNED | Like beachcombers or surfers |
| SEASHELLS | Featuring in the rocaille rockwork of grottoes and later in aspects of rococo, the "coquilles" or "conchiglie" of angel wings, cockles, jingles, periwinkles, volutes, wentletraps and other marine moll |
| SHELL | Exoskeleton of a snail; or, part of tortoise consisting of a carapace, plastron and bridge (5) |
| SONIC | Video-game character who collects the Chaos Emeralds |
| FIELDSMAN | Cricketer who collects the ball once it is struck (9) |
| FIELDSMEN | Cricketer who collects the ball once it is struck (9) |
| POLYP | Tiny organism - coral reefs are formed of the calcium carbonate exoskeleton of many such creatures |
| SHELLS | Exterior carapaces of molluscs including periwinkles, cockles, clams, oysters, scallops and wentletraps (6) |
| ROUNDSUP | How the cowboy collects the cows till the cows come home (6,2) |
| MOLLY | - Malone; purveyor of cockles and mussels in an Irish folk song of an same name which is recognised as Dublin's unofficial anthem (5) |
| GASTROPOD | To what class of mollusc do slugs, snails, cowries and whelks belong? (9) |
| PRESERVE | Dr Strabismus (Whom God ____) of Utrecht was a character in the Daily Express Beachcomber column |
| GATHERS | It's the way he collects the rags that counts (7) |
| ROCKPOOLS | Puddles of trapped sea water forming mini ecosystems or habitats for limpets, periwinkles, anemones, starfish and other marine life (4,5) |
| OYSTERS | Molluscs in a phylum with clams, mussels, cockles and periwinkles (7) |