| LINDAUER | Artist, born in Western Bohem: in 1839, best known for his studies of Maori (8) |
| BATWOMAN | Superheroine born in western Arabian state (8) |
| CARLJUNG | Swiss psychologist (1875-1961) known for his studies of the human psyche (4,4) |
| LANDSEER | English painter noted for his studies of animals (8) |
| HANSSELYE | Endocrinologist known for his studies of the effects of stress on the human body (4,5) |
| STUBBS | George, painter known for his studies of horses such as 1762's Whistlejacket (6) |
| PAVLOV | Russian physiologist, known chiefly for his studies of conditioned reflex behaviour, especially in dogs (6) |
| MEADE | English economist noted for his studies of international trade and finance (born in 1907). |
| SELYE | Endocrinologist noted for his studies of the effects of stress on the body (5) |
| WAITANGI | The Treaty of ___, 6 February 1840, between Britain and a number of Maori tribes, which effectively founded New Zealand (8) |
| TUKUTUKU | What hand-woven latticework wall panels are a traditional form of Maori art? (8) |
| GALVANI | 18th-century Italian physician and physicist noted for his studies of the effects of electricity on animal nerves and muscles (7) |
| LUIGI | 18th-century Italian physician and physicist noted for his studies of the effects of electricity on animal nerves and muscles (5) |
| GALEN | Pre-eminent physician of ancient Greece after Hippocrates, noted especially for his studies of anatomy (129-c. 216 AD) (5) |
| DEGAS | Edgar, French impressionist painter noted for his studies of horse racing and ballet dancers (5) |
| REDHEADS | Edition initially hailed in studies of Titian subjects (8) |
| INSPECTS | Studies of 21 types of creature found outside Portugal (8) |
| CANALETTO | Italian painter noted for his studies of Venice (9) |
| STRUVE | Otto -; Russian-born US astronomer noted for his studies in stellar spectroscopy and discovery of ionised interstellar hydrogen (6) |
| EADWEARDMUYBRIDGE | English-born US photographer noted for his pioneering high-speed photographic studies of animals and people in motion |