| LEAF | A folio of a book; a botanical structure known in its embryonic form as a cotyledon; or, gold in the form of foil, used in gilding (4) |
| PAGE | One side of a leaf or a folio of a book; or, historically, a boy in training for knighthood (4) |
| ROOT | A structure known in its edible form as a carrot, parsnip, radish, swede or turnip, for example (4) |
| GERMINAL | Zola novel in its embryonic stage (8) |
| FILLET | From "thread", a word for a narrow ribbon tied round the head originally, later a boned side of fish or slice of meat; a decorative line impressed on the cover of a book; a narrow piece of wood; or, a |
| TOWER | Bell -; structure known in Italian as a campanile (5) |
| DAF | Hebrew for page; a folio of Talmud |
| OVULE | From Latin for "egg", a botanical structure that develops into a seed (5) |
| SPINE | Heartwood of a tree; a quill of a hedgehog; a bound edge of a book; a series of vertebrae; or, courage (5) |
| FLOWER | General word for a botanical structure essential for pollinators including bees and moths (6) |
| BUD | Bloom in embryonic form (3) |
| BLASTULA | Rudimentary embryonic form |
| JACKET | Skin of a potato; dust cover of a book; a record sleeve; or, an outer garment (6) |
| ASYOULIKEIT | Pastoral comedy originally published in the First Folio of 1623 (2,3,4,2) |
| SEEDLEAF | Cotyledon |
| LEWISIA | - - - cotyledon, evergreen perennial commonly called cliff maids (7) |
| WATER | The brilliance, lustre and purity of a diamond, such as one in the first degree, considered flawless; a wavy sheen on moire silk or taffeta; or, a liquid compound known in its solid or gaseous state a |
| VERSO | From "turned leaf", the back of a folio or sheet; a left-hand page of an open book; or, the rear of something, such as a coin, medal or painting (5) |
| SCULPTURE | Work of art such as a bronze, known in its preliminary wax or clay form as a maquette (9) |
| SWATCH | Small sample of carpet, cloth or wallpaper; a number of said testers bound as a book; a computer's palette of active colours; or, a spot of ink for matching during printing (6) |
