| ASHDOWN | - Forest; area of heathland in Sussex that inspired A. A. Milne's fictional Hundred Acre Wood (7) |
| GRAHAME | Author whose classic tale The Wind in the Willows inspired A A Milne's play Toad of Toad Hall (7) |
| ACRE | Five Hundred - Wood; part of Ashdown Forest in East Sussex that inspired A. A. Milne (4) |
| MILNE | Children's author who wrote about the fictional Hundred Acre Wood (5) |
| BODMIN | --- Moor is a barren heathland in Cornwall, England (6) |
| BODMINMOOR | Barren heathland in Cornwall, England (6,4) |
| ADUR | River in Sussex that reaches the English Channel at Shoreham-by-Sea |
| GLYNDEBOURNE | Opera venue in Sussex that hosts an annual festival (12) |
| ARUNDEL | Market town in West Sussex that has a medieval castle and a Roman Catholic cathedral (7) |
| WEALDEN | Local government district in East Sussex that includes the market town of Uckfield (7) |
| CALLUNA | From "beautify, sweep clean", the genus of heathland heathers or ling with masses of purple or rose-pink bell-shaped blooms on stems traditionally bunched as brooms (7) |
| ARGONNE | Forest area half destroyed and dead in which knight is buried (7) |
| ALAMEIN | A.A. Milne's work achieved fame in World War II (7) |
| WOOZLES | Imaginary honey-stealing creatures in A. A. Milne's book Winnie-the-Pooh, often associated with heffalumps (7) |
| EPPING | --- Forest, area of ancient woodland in Southern England, a former royal hunting ground (6) |
| ARDEN | Old forest area of Warwickshire ... in hard enclosure (5) |
| SILVERSTUDDEDBLUE | Butterfly, once common in the UK, in decline due to loss of heathland |
| HOVE | Seaside resort in East Sussex that achieved city status with Brighton in 2000 |
| RYE | Ancient Town in East Sussex that joined the confederation of Cinque Ports (3) |
| LIVETOLEAD | Show presented by the Duke and Duchess of Sussex that debuted in 2022 |
